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Toll/interleukin-1 receptor domain-containing adapter protein; TIR domain-containing adapter protein; adaptor protein Wyatt; MyD88 adapter-like protein (TIRAP, MAL)

Function: - adapter involved in TLR4 signaling - innate immune response - acts via IRAK2 & TRAF-6 - activation of NF-kappa-B, MAPK1, MAPK3 & JNK - cytokine secretion - inflammatory response - positively regulates production of TNF-alpha & IL-6 - phosphorylated by IRAK1 & IRAK4 - also phosphorylated by BTK - polyubiquitination follows phosphorylation by BTK & leads to TIRAP degradation - forms homodimers & heterodimers with MyD88 - binds to TLR4 via their TIR domain - binds to BMX, PKR & TBK1 - does not interact with IRAK1, nor TLR9 Structure: - homodimer - contains 1 TIR domain Compartment: cytoplasm Alternative splicing: named isoforms=3 Expression: - expressed in liver, kidney, spleen, skeletal muscle, heart > peripheral blood leukocytes, lung, placenta, small intestine, thymus, colon, brain Polymorphism: - genetic variations in TIRAP may influence susceptibility or resistance to invasive pneumococcal disease [2], malaria [3], & tuberculosis [4] - genetic variations in TIRAP influence susceptibility or resistance to bacterial invasion of the blood & define the bacteremia susceptibility locus 1 (BACTS1) [5]

Related

interleukin-1 receptor (IL1R) Toll-like receptor 4; hToll; CD284 (TLR4)

General

glycoprotein secreted protein

Properties

SIZE: entity length = 221 aa MW = 24 kD COMPARTMENT: cytoplasm MOTIF: glycosylation site TIR domain {84-221} MOTIF: cysteine residue {C89} MODIFICATION: cysteine residue {C134} cysteine residue {C134} MODIFICATION: cysteine residue {C89} cysteine residue {C142} MODIFICATION: cysteine residue {C174} cysteine residue {C174} MODIFICATION: cysteine residue {C142}

Database Correlations

OMIM correlations MORBIDMAP 606252 UniProt P58753 Pfam PF13676 Entrez Gene 114609 Kegg hsa:114609

References

  1. UniProt :accession P58753
  2. OMIM :accession 610799
  3. OMIM :accession 611162
  4. OMIM :accession 607948
  5. OMIM :accession 614382
  6. SeattleSNPs http://pga.gs.washington.edu/data/tirap/