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protein ajuba (AJUBA, JUB)

Function: - component of cellular adhesives complexes - contributes to cell fate determination & regulates cell proliferation & differentiation - regulation of actin cytoskeleton dynamics & cell migration - role in linking &/or strengthening of epithelia cell-cell junctions in part by linking adhesive receptors to the actin cytoskeleton - role in signal transduction from cell adhesion sites to the nucleus - role in regulation of the kinase activity of AURKA/Aurora-A for mitotic commitment - component of the IL-1 signaling pathway modulating IL-1-induced NF-kappa-B activation by influencing the assembly & activity of PRKCZ/SQSTM1/TRAF6 multiprotein signaling complex - can function as adapter molecule in formation of large multiprotein complexes that form on DNA & that influence transcription - interacts with AURKA/Aurora-A during mitosis & both proteins are phosphorylated as they form a complex. - interacts with CTNNA1/alpha-catenin & with F-actin - interacts with LATS2 during mitosis & this complex regulates organization of the spindle apparatus through recruitment of gamma-tubulin to the centrosome - interacts with GRB2 & PI51A (putative) - forms a complex with SQSTM1, PRKCZ & TRAF6 - interacts with GLT1 (putative) - phosphorylated by LATS2 during mitosis - phosphorylated by AURKA Structure: - LIM region interacts with CTNNA1 - preLIM region binds directly actin filaments - LIM-2 & LIM-3 domains mediate the interaction with the N-terminal region of AURKA - association between LATS2 & JUB requires 2nd LIM domain of JUB - belongs to the zyxin/ajuba family - contains 3 LIM Zn+2-binding domains Compartment: - cytoplasm, cell membrane, cell junction, nucleus, centrosome - shuttles between the cytoplasm & the nucleus - localizes on centrosomes during G2-M phase - preferentially co-localizes with cadherin-adhesive complexes at sites of cell-cell contacts Notes: 'Ajuba' means 'curiosity' in Urdu, an Indian dialect

General

cytoskeletal protein phosphoprotein

Properties

SIZE: entity length = 538 aa MW = 57 kD COMPARTMENT: cytoplasm cell nucleus MOTIF: PreLIM {1-335} MOTIF: Ser phosphorylation site {S119} Ser phosphorylation site {S263} nuclear translocation signal {280-288} LIM domain {336-397} MOTIF: cysteine residue (6) histidine residue Zn+2-binding site LIM domain {401-461} MOTIF: cysteine residue (6) histidine residue Zn+2-binding site LIM domain {462-530} MOTIF: cysteine residue (6) histidine residue Zn+2-binding site

Database Correlations

UniProt Q96IF1 Pfam PF00412

References

UniProt :accession Q96IF1