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interleukin-12 receptor beta-2 chain; IL-12 receptor beta-2; IL-12R-beta2 (IL12RB2)

Function: - with I12RB1, forms functional high affinity IL12 receptor - signaling component coupling to the JAK2/STAT4 pathway - promotes the proliferation of T-cells as well as NK cells - induces the promotion of T-cells towards the Th1 phenotype by strongly enhancing IFN-gamma production - Il12RB2 binds JAK2 (via its N-terminal) through a membrane-proximal region of the cytoplasmic domain - interaction, in vitro & in vivo, with SOCS3 (via its SH2 domain) inhibits the STAT4-mediated activation - binds STAT4 through a membrane-distal C-terminal region - on IL12 binding, phosphorylated on C-terminal Tyr by JAK2 - phosphorylation on Tyr-800 is required for STAT4 binding & activation, & for SOCS3 binding Structure: - the WSXWS motif appears to be necessary for proper protein folding, efficient intracellular transport & cell-surface receptor binding - the box 1 motif is required for JAK interaction &/or activation - belongs to the type I cytokine receptor family, type 2 subfamily - contains 5 fibronectin F3 modules Compartment: membrane Alternative splicing: named isoforms=2 Expression: - isoform 2 is expressed at similar levels in both naive & activated T-cells - maximum levels in Th1 cells between day 3 & day 8 of activation - in vitro, up-regulated by interferon alpha Polymorphism: - heterozygotic variants Gly-313 & Arg-720 are associated with atopy

General

glycoprotein membrane protein protein subunit

Properties

SIZE: entity length = 862 aa MW = 97 kD COMPARTMENT: cellular membrane MOTIF: signal sequence {1-23} N-glycosylation site {N48} fibronectin type III domain or F3 module {124-218} MOTIF: N-glycosylation site {N129} N-glycosylation site {N166} N-glycosylation site {N195} serine-rich region {208-211} MOTIF: serine residue (SEVERAL) fibronectin type III domain or F3 module {224-316} MOTIF: N-glycosylation site {N271} WSXWS motif {305-309} fibronectin type III domain or F3 module {317-415} MOTIF: N-glycosylation site {N347} N-glycosylation site {N376} fibronectin type III domain or F3 module {420-517} MOTIF: N-glycosylation site {N480} fibronectin type III domain or F3 module {521-617} transmembrane domain {623-643} box1 motif SITE: 662-670 FOR-BINDING-OF: Janus kinase (JAK) proline-rich region SITE: 750-753 MOTIF: proline residue (SEVERAL) STAT4 binding {796-801} MOTIF: Tyr phosphorylation site {Y800}

Database Correlations

OMIM 601642 UniProt Q99665 PFAM correlations Entrez Gene 3595 Kegg hsa:3595

References

  1. UniProt :accession Q99665
  2. SeattleSNPs http://pga.gs.washington.edu/data/il12rb2/

Component-of

interleukin-12 receptor