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A disintegrin & metalloproteinase domain 20 (ADAM20)

Function: - may be involved in sperm maturation &/or fertilization - no obvious cleavage site for furin endopeptidase, suggesting that the proteolytic processing is regulated Cofactor: binds 1 Zn+2 per subunit (putative) Structure: - a tripeptide motif (VGE) within disintegrin-like domain could be involved in the binding to egg integrin receptor & thus could mediate sperm/egg binding - cysteine-rich domain encodes putative cell-fusion peptides, which could be involved in sperm-egg fusion - conserved Cys present in the cysteine-switch motif binds the catalytic Zn+2, thus inhibiting the enzyme - dissociation of Cys from Zn+2 upon the activation- peptide release activates the enzyme - contains 1 disintegrin domain - contains 1 EGF-like domain - contains 1 peptidase M12B domain Compartment: membrane Expression: testis specific Notes: - may be the functional equivalent of ADAM1 (fertilin alpha) which is a pseudogene in human

General

ADAM (A disintegrin & metalloproteinase domain); MDC (metalloproteinase, disintegrin, cysteine-rich) protein glycoprotein testis-specific protein

Properties

SIZE: MW = 82 kD entity length = 726 aa COMPARTMENT: cellular compartment CELL: unspecified cell type WITHIN: testis MOTIF: N-glycosylation site {N191} metalloprotease domain {207-384} MOTIF: N-glycosylation site {N226} cysteine residue {C317} MODIFICATION: cysteine residue {C394} active site SITE: E343 cysteine residue {C357} MODIFICATION: cysteine residue {C379} N-glycosylation site {N378} cysteine residue {C379} MODIFICATION: cysteine residue {C357} cysteine residue {C394} MODIFICATION: cysteine residue {C317} disintegrin domain {407-493} MOTIF: N-glycosylation site {N438} cysteine residue {C465} MODIFICATION: cysteine residue {C478} cysteine residue {C478} MODIFICATION: cysteine residue {C465} N-glycosylation site {N479} cysteine-rich region {494-634} MOTIF: N-glycosylation site {N587} EGF domain {635-663} MOTIF: cysteine residue {C635} MODIFICATION: cysteine residue {C646} cysteine residue {C640} MODIFICATION: cysteine residue {C652} cysteine residue {C646} MODIFICATION: cysteine residue {C635} cysteine residue {C652} MODIFICATION: cysteine residue {C640} cysteine residue {C654} MODIFICATION: cysteine residue {C663} cysteine residue {C663} MODIFICATION: cysteine residue {C654} transmembrane domain {694-714}

Database Correlations

OMIM 603712 UniProt O43506 PFAM correlations Entrez Gene 8748 Kegg hsa:8748

References

UniProt :accession O43506