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breakpoint cluster region protein P160 (p160BCR)

Protein product of BCR-gene (breakpoint cluster region) gene that forms a chimeric gene with abl in the Philadelphia chromosome associated with chronic myelogenous leukemia (CML). Function: - GTPase-activating protein for RAC1 & CDC42 - promotes exchange of RAC or CDC42-bound GDP by GTP, thereby activating them - displays serine/threonine kinase activity - interacts with PDZK1 - may interact with CCPG1 - autophosphorylated ATP + a protein ADP + a phosphoprotein Structure: - homotetramer - region involved in binding to ABL1 SH2-domain is rich in Ser & needs to be Ser/Thr phosphorylated prior to SH2 binding; this region is essential for the activation of the ABL1 tyrosine kinase & transforming potential of the chimeric BCR-ABL oncogene - DH domain is involved in interaction with CCPG1 - contains 1 C2 domain contains 1 DH (DBL-homology) domain contains 1 PH domain contains 1 Rho-GAP domain Alternative splicing: named isoforms=2 Pathology: - chromosomal translocation t(9;22)(q34.1;q11) involving BCR with ABL1 is a cause of chronic myeloid leukemia translocation produces a BCR-ABL found also in acute myeloid leukemia (AML) & acute lymphoblastic leukemia (ALL))

Related

guanine nucleotide exchange factor (GDP/GTP dissociation stimulator, guanine nucleotide-releasing factor/protein, GNEF, GNDS, GDS)

Specific

p210 bcr/abl

General

oligomerizing protein rho GTPase-activating protein (rho-GAP) serine/threonine kinase

Properties

SIZE: MW = 160 kD STATE: active state MOTIF: Kinase {1-426} MOTIF: Tyr phosphorylation site {Y177} Binding to ABL SH2 {197-385} Thr phosphorylation site {T310} Breakpoint {426-427} Ser phosphorylation site {S459} Ser phosphorylation site {S463} Dbl homology domain {498-691} MOTIF: Tyr phosphorylation site {Y591} Tyr phosphorylation site {Y644} PH domain {708-866} MOTIF: leucine residue {824-827} (4) C2 domain {870-1002} MOTIF: binding site FOR-BINDING-OF: phospholipid Ca+2-binding site rho-GAP domain NAME: rho-GAP domain SITE: 1054-1248 Ser phosphorylation site {S1264}

Database Correlations

OMIM correlations UniProt P11274 PFAM correlations Entrez Gene 613

References

  1. Maru Y, Witte ON. The BCR gene encodes a novel serine/threonine kinase activity within a single exon. Cell. 1991 Nov 1;67(3):459-68. PMID: 1657398
  2. Musacchio A, Gibson T, Rice P, Thompson J, Saraste M. The PH domain: a common piece in the structural patchwork of signalling proteins. Trends Biochem Sci. 1993 Sep;18(9):343-8. Review. PMID: 8236453
  3. Atlas of Genetics & cytogenetics in oncology & Haematology http://atlasgeneticsoncology.org/Genes/BCR.html