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cyclin-dependent kinase-1 (CDK1), cell division control protein-2 (cdc2), p34 protein kinase or p34cdc2

Founding member of the cyclin-dependent kinase family. Function: - CDK1 is a component of MPF (Maturation Promoting Factor) & of CTD kinase - role in the control of the eukaryotic cell cycle - required in higher cells for entry into S-phase & mitosis - p34 is a component of the kinase complex that phosphorylates the repetitive C-terminus of RNA polymerase 2 - forms a stable but non-covalent complex with a regulatory subunit & with a cyclin - interacts with DLG7 - isoform 2 is unable to complex with cyclin B1 & also fails to bind to the CDK inhibitor p21 - interacts with catalytically active CCNB1 & RALBP1 during mitosis to form an endocytotic complex during interphase - phosphorylation at Thr-14 or Tyr-15 inactivates the enzyme - phosphorylation at Thr-161 activates it - component of MPF (Maturation Promoting Factor) - component of CTD kinase. Structure: - belongs to the protein kinase superfamily, CMGC Ser/Thr protein kinase family, CDC2/CDKX subfamily - contains 1 protein kinase domain Compartment: nucleus (putative) Alternative splicing: named isoforms=2 Pathology: found in breast cancer tissues Comparative biology: - Yeast homologs are cdc2 (S. cerevisiae), cdc28 (S. pombe), cdc2-kinase - Thr-14 phosphorylation site is not present in yeast homologs

General

cyclin-dependent kinase (CDK)

Properties

SIZE: MW = 34 kD STATE: Inactive MOTIF: kinase domain SITE: 4-287 MOTIF: ATP-binding site NAME: ATP-binding site SITE: 10-18 Thr phosphorylation site {T14} Tyr phosphorylation site {Y15} Tyr phosphorylation site {Y19} ATP-binding site NAME: ATP-binding site SITE: 33-33 aspartate residue {D128} Thr phosphorylation site {T161}

Database Correlations

OMIM 116940 UniProt P06493 Pfam PF00069 Entrez Gene 983 Kegg hsa:983 ENZYME correlations

References

  1. Gould KL, Moreno S, Tonks NK, Nurse P. Complementation of the mitotic activator, p80cdc25, by a human protein-tyrosine phosphatase. Science. 1990 Dec 14;250(4987):1573-6. PMID: 1703321
  2. De Bondt HL, Rosenblatt J, Jancarik J, Jones HD, Morgan DO, Kim SH. Crystal structure of cyclin-dependent kinase 2. Nature. 1993 Jun 17;363(6430):595-602. PMID: 8510751 {CDK2 crystal}
  3. Morgan DO. Principles of CDK regulation. Nature. 1995 Mar 9;374(6518):131-4. Review. PMID: 7877684

Component-of

CDK1/cyclin B complex or maturation promoting factor (MPF)